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Plant cells are surrounded by a tough extracellular matrix in the form of a cell wall, which is responsible for many of the unique features of a plant's life style. The cell wall is composed of a network of cellulose microfibrils and cross-linking glycans embedded in a highly cross-linked matrix of pectin polysaccharides. In secondary cell walls, lignin may be deposited. A cortical array of microtubules can determine the orientation of newly deposited cellulose microfibrils, which in turn determines directional cell expansion and therefore the final shape of the cell and, ultimately, of the plant as a whole.