Your friend Mabel studies wild populations of bison in Yellowstone National Park. She notes that the mean horn length is 30 cm with a variance of 5 cm. She knows that bison horns are an important defense weapon and is curious if evolution might select for longer horns. After intense study of the bison population, Mabel determines the variance components to be: VA=0.1, VD=0.2, VI=0.3, VG=0.6, VE=3.4, VGE=1.0. In your informed opinion, how well will horn length respond to selection for longer horns in this population?

searching efficiency (a)

Explanation:

The other word for Lotka-Volterra is predator-prey in which they have a combination of the differential equation which consists of nonlinear and should be the first ordered sequence.

It is used to tell the biological system changes where the two species interacted with each other. The one is the predator and the other one is prey.

Now according to the question the rate at which the  predator-prey change is searching efficiency (a) as it shows the changes of predator species with the respect that how quick they determine, attack it and consumed it